PASADENA, Calif. (AP) Fresno State's Jake Haener was able to overcome a crushing hit to his right hip to lead the Bulldogs to an upset victory over No. 13 UCLA.

Haener threw for 455 yards, including the go-ahead touchdown pass to Jalen Cropper with 14 seconds remaining, as the Bulldogs beat the Bruins 40-37 Saturday night at the Rose Bowl.

''We know he's tough, right. We've seen that over the course of last year and this year. He's tough,'' Fresno State coach Kalen DeBoer said. ''When the pressure was great, he was at his best and that's what's awesome.''

Haener had to be at his best in a game that saw four lead changes over the final 7:27 after the Bulldogs had a 23-10 lead at halftime.

Fresno State finished the job against a Pac-12 program after leading then-No. 11 Oregon in the fourth quarter two weeks ago before losing.

''I mean, we just made one more play, right? We made one more play,'' DeBoer said. ''Fifty-four seconds that I just think about what our guys are capable of doing, it didn't feel like it was too big of a feat at all.''

Haener's 19-yard strike to Erik Brooks put Fresno State back in front 33-30 with 2:55 remaining, but he was wincing in pain on the ground after taking numerous hits throughout the night.

After UCLA regained the lead with 54 seconds left on Dorian Thompson-Robinson's 15-yard pass to Kyle Phillips, a limping Haener directed a six-play, 75-yard drive that gave the Bulldogs (3-1) their fourth straight win over the Bruins.

''He was a pretty mobile quarterback. We struggled to keep him in the pocket,'' UCLA cornerback Qwuantrezz Knight said.

Cropper had three receptions for 50 yards on the drive, including the TD. Josh Kelly also had a 26-yard catch that put the ball at the UCLA 12 with 23 seconds remaining.

''Yeah, I'm not gonna lie, I was a little tired,'' Cropper said. ''But that last play, I saw the corner drop. And just knowing how Jake trusts me throwing that back shoulder into the corner of the end zone, the feeling's so surreal. I can't really describe it in words.''

Haener completed 38 of 53 passes and tossed two touchdowns. Cropper had 14 receptions for 141 yards and Kelly added eight catches for 120 yards. Ronnie Rivers rushed for 136 yards and two touchdowns.

''I think the quarterback is a special player,'' UCLA coach Chip Kelly said. ''We threw everything at him. We blitzed him, played cover zero, rushed three and dropped everyone else back into coverage. I think we threw the entire kitchen sink at him and he handled everything.''

UCLA was able to rally back due to a pair of Bulldogs turnovers. Fresno State led 26-17, had the ball at the UCLA 9 and were looking to put the game away with 9:23 remaining when Carl Jones stripped Haener of the ball. After it rolled 34 yards and a mad scramble ensued, Myles Jackson recovered it at the UCLA 43.

Three plays later, Thompson-Robinson delivered a perfect pass to Phillips, who eluded Bralyn Lux's coverage, for a 38-yard score to bring UCLA within 26-23.

The Bulldogs wouldn't have it long on the ensuing possession when Kelly fumbled after a 13-yard completion. UCLA's Jordan Genmark Heath recovered and returned it 20 yards to the Bulldogs 18.

UCLA took its first lead since the first quarter when Zach Charbonnet capped off a three-play drive with a 3-yard TD up the middle to put the Bruins up 30-26.

Thompson-Robinson passed for 278 yards and three touchdowns while Phillips had seven receptions for 113 yards and two TDs. Charbonnet ran for two touchdowns for the Bruins (2-1), who were ranked for the first time in four years.

Thompson-Robinson passed for 278 yards and three touchdowns while Phillips had seven receptions for 113 yards and two TDs. Charbonnet ran for two touchdowns for the Bruins (2-1), who were ranked for the first time in four years.

''We definitely made it tough on ourselves. We made mistakes and bit ourselves in the foot too many times,'' Thompson-Robinson said.

Rivers scored both of his touchdowns in the first half. He opened the scoring with a 1-yard run up the middle with 5:54 remaining in the first quarter to give the Bulldogs a 6-0 lead. The 2-point conversion was no good after holder Carson King's pass was incomplete.

Rivers then broke a pair of tackles en route to a 36-yard score up the right sideline midway through the second quarter to extend Fresno State's lead to 20-10.

THE TAKEAWAY

Fresno State: Haener has thrown for over 300 yards in three of the Bulldogs' first four games and leads the nation with 1,464 passing yards.

UCLA: The Bruins need to clean up their pass defense and penalties if they hope to bounce back. They had two pre-snap penalties in punt coverage on fourth down that allowed the Bulldogs to keep the ball.

POLL IMPLICATIONS

UCLA might remain in the polls after showing the mettle to rally back, but it was a bad week for the Pac-12. The five teams in the South Division that had non-conference games were favored and lost.

UP NEXT

Fresno State begins its Mountain West schedule on Friday against UNLV.

UCLA travels to Stanford on Saturday to begin Pac-12 play.

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